Bontrager's 29-2 Expert is specifically engineered for 29ers with a tread design that is perfect for loose-over-hardpack or loamy trails. A shallow center tread rolls fast, then you've got scooped edges that dig into loose terrain and longer shoulder blocks for stability in corners. And the folding bead keeps the weight low.
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The middle grips wore off with in the first hundred miles. They were useless in wet conditions. Using them to climb hills that had any kind of rock or roots was pointless, unless you are looking for a workout. Constantly slipped going up rocky, or wet, or rooty hills. Once worn down they were basically rode bike tires that used a lot of energy because of the width.